Published January 24, 2023. Updated February 8, 2024. Open access. Peer-reviewed.

Gallery ❯

Common Stream-Lizard (Potamites ecpleopus)

Reptiles of Ecuador | Sauria | Gymnophthalmidae | Potamites ecpleopus

English common name: Common Stream-Lizard.

Spanish common name: Lagartija ribereña común.

Recognition: ♂♂ 17.6 cmMaximum distance from the snout to the tip of the tail. Snout–vent length=8.4 cm. ♀♀ 16.8 cmMaximum distance from the snout to the tip of the tail. Snout–vent length=7.1 cm..1,2 Stream-Lizards of the genus Potamites are easily distinguishable from other leaf-litter lizards in their area of distribution by having heterogeneous dorsal scales and two longitudinal crests on the tail.3 The Common Stream-Lizard (P. ecpleopus) can be confused with its congener P. strangulatus and with species of the genus Gelanesaurus, from which it differs by having well-developed caudal ridges and enlarged tubercles around the neck, flanks, tail, and upper surface of the extremities, all of which give the lizard a crocodile-like appearance.15 There is much geographic variation in P. ecpleopus.4,6 In Ecuadorian populations, adult males have alternating black and white bars on the lips and edges of the chin, a dark throat coloration, and black ocelli with a cream center along the flanks. In females, the labial bars are dark brown instead of black, the throat is light gray, and there is a single ocellus on the insertion of the arm.1,7 In adults, the dorsal coloration is tan to dull brown, whereas in juveniles the back is pinkish tan.1

Figure showing variation among individuals of Potamites ecpleopus

Figure 1: Individuals of Potamites ecpleopus: La Bonita, Sucumbíos province, Ecuador (); Palmarí Reserve, Amazonas state, Brazil (); Tzarentza, Pastaza province, Ecuador (); Río Bigai Reserve, Orellana province, Ecuador (). j=juvenile.

Natural history: In Ecuador, Potamites ecpleopus is considered a common lizard that occurs in high densities.7,8 It inhabits old-growth rainforests both in flooded and terra-firme areas,610 but also in disturbed areas such as pastures11 and habitats affected by mining or logging.2,8 These lizards are diurnal, semi-aquatic, and strongly associated with bodies of water.4,9,12 They are good swimmers; in water, they propel themselves by means of sinuous movements of the body with help of the tail.4,13 They also forage among leaf-litter, rocks, and logs along or around swampy areas, creeks, streams, and waterfalls,7,14,15 usually partially submerged.1,12 On occasion, these reptiles perch on vegetation up to almost 1 m above the ground or sit exposed on logs and rocks, basking in areas with filtered light.16 However, they prefer humid and shady areas within the forest and do not require direct sunlight to thermoregulate, being active during cloudy and rainy days, even in cold water.1,9,17 At night, they roost on vegetation or among sticks and leaf-litter near streams.8,13 They move slower than other co-occurring lizards and their body temperature is lower,6,7 between 24–30°C, and directly influenced by the air or ground temperature.14

Individuals of Potamites ecpleopus return to the same point on consecutive days to feed.12 Their diet is considered generalist and includes a wide variety of invertebrates ranging from earthworms to snails, but the most important prey items are crickets, ants, and insect larvae.6,7 In smaller quantities, it also includes tadpoles17 and lizards such as Anolis trachyderma, Loxopholis parietalis, and even other P. ecpleopus.1 There are records of snakes (Drymoluber dichrous,1 Helicops angulatus,9 Oxyrhopus leucomelas,8 and Bothrocophias hyoprora18) preying upon individuals of P. ecpleopus. When threatened, these lizards run towards roots and leaf-litter, into holes in the banks,9 or directly to the water4,7 submerging to the bottom debris.1,6,14 If captured, they usually wriggle and bite fiercely, but can also shed the tail.1,8 Potamites ecpleopus is an oviparous species with a continuous reproductive cycle.9,19 The clutch size is two eggs measuring around 0.5 x 1.4 cm.2 Communal nests of up to 5 eggs have been reported under large, rotten logs near bodies of water.4 Hatchlings measure 2.1–2.4 cm in SVL.9

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Conservation: Least Concern Believed to be safe from extinction given current circumstances..20 Potamites ecpleopus is listed in this category because it has a large distribution, occurs in protected areas, and has high population densities. Thus, the species is considered to be facing no major immediate extinction threats.20 Although deforestation is affecting some populations, this is not considered a widespread threat to the species as a whole.20 However, P. ecpleopus has been shown to be a species complex21 and some populations are under greater threat from habitat loss.

Distribution: Potamites ecpleopus is widely distributed throughout the Amazon basin of Bolivia, Brazil, Colombia, Ecuador (Fig. 2), and Perú.2 It also occurs along the adjacent Andean foothills.

Distribution of Potamites ecpleopus in Ecuador

Figure 2: Distribution of Potamites ecpleopus in Ecuador. See Appendix 1 for a complete list of the presence localities included in the map.

Etymology: The generic name Potamites, which is a Greek noun meaning “water finder,” refers to the semi-aquatic habits of lizards in this genus.3 The specific epithet ecpleopus refers to the similarity between the head of this species and those of the tribe Ecpleopini.22

See it in the wild: Common Stream-Lizards can be observed with almost complete certainty in forested streams throughout the species’ area of distribution in Ecuador. These reptiles are particularly common in Yasuní National Park, Sumaco National Park, and Cuyabeno Wildlife Reserve. Individuals can be found by removing leaf-litter or logs along streams or flooded areas within the forest

Special thanks to Daniel Sheehan for symbolically adopting the Common Stream-Lizard and helping bring the Reptiles of Ecuador book project to life.

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Authors: Amanda QuezadaaAffiliation: Tropical Herping (TH), Quito, Ecuador.,bAffiliation: Laboratorio de Herpetología, Universidad del Azuay, Cuenca, Ecuador. and Alejandro ArteagacAffiliation: Khamai Foundation, Quito, Ecuador.

Academic reviewer: Jeffrey D CamperdAffiliation: Department of Biology, Francis Marion University, Florence, USA.

Photographers: Jose VieiraaAffiliation: Tropical Herping (TH), Quito, Ecuador.,eAffiliation: ExSitu, Quito, Ecuador. and Sebastián Di DoménicofAffiliation: Keeping Nature, Bogotá, Colombia.

How to cite? Quezada A, Arteaga A (2024) Common Stream-Lizard (Potamites ecpleopus). In: Arteaga A, Bustamante L, Vieira J (Eds) Reptiles of Ecuador: Life in the middle of the world. Available from: DOI: 10.47051/EMMQ9145

Literature cited:

  1. Duellman WE (1978) The biology of an equatorial herpetofauna in Amazonian Ecuador. Publications of the Museum of Natural History, University of Kansas 65: 1–352.
  2. Avila-Pires TCS (1995) Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen 299: 1–706.
  3. Doan TM, Castoe TA (2005) Phylogenetic taxonomy of the Cercosaurini (Squamata: Gymnophthalmidae), with new genera for species of Neusticurus and Proctoporus. Zoological Journal of the Linnean Society 143: 405–416. DOI: 10.1111/j.1096-3642.2005.00145.x
  4. Uzzell TM (1966) Teid Lizards of the genus Neusticurus (Reptilia, Sauria). Bulletin of the American Museum of Natural History 132: 279–327.
  5. Altamirano-Benavides M, Zaher H, Lobo L, Grazziotin FG, Sales Nunes PM, Rodrigues MT (2013) A new species of lizard genus Potamites from Ecuador (Squamata, Gymnophthalmidae). Zootaxa 3717: 345–358. DOI: 10.11646/zootaxa.3717.3.4
  6. Vitt LJ, Avila-Pires TCS (1998) Ecology of two sympatric species of Neusticurus (Sauria: Gymnophthalmidae) in the Western Amazon of Brazil. Copeia 1998: 570–582. DOI: 10.2307/1447787
  7. Vitt LJ, De la Torre S (1996) A research guide to the lizards of Cuyabeno. Pontificia Universidad Católica del Ecuador, Quito, 165 pp.
  8. Field notes, Reptiles of Ecuador book project.
  9. Dixon JR, Soini P (1986) The reptiles of the upper Amazon Basin, Iquitos region, Peru. Milwaukee Public Museum, Milwaukee, 154 pp.
  10. Catenazzi A, Lehr E, von May R (2013) The amphibians and reptiles of Manu National Park and its buffer zone, Amazon basin and eastern slopes of the Andes, Peru. Biota Neotropica 13: 269–283. DOI: 10.1590/S1676-06032013000400024
  11. Camper JD, Torres-Carvajal O, Ron SR, Nilsson J, Arteaga A, Knowles TW, Arbogast BS (2021) Amphibians and reptiles of Wildsumaco Wildlife Sanctuary, Napo Province, Ecuador. Check List 17: 729–751.
  12. Rocha CFD (1991) Habitat utilization and feeding habits of Neusticurus ecpleopus in a Brazilian tropical rainforest. Herpetological Review 22: 40–42.
  13. Whitworth A, Beirne C (2011) Reptiles of the Yachana Reserve. Global Vision International, Exeter, 130 pp.
  14. Fitch HS (1968) Temperature and behavior of some equatorial lizards. Herpetologica 24: 35–38.
  15. da Fonseca WL, da Silva JD, Abegg AD, da Rosa CM, Bernarde PS (2019) Herpetofauna of Porto Walter and surrounding areas, southwest Amazonia, Brazil. Herpetology Notes 12: 91–107.
  16. Vitt LJ, Zani PA (1996) Organization of a taxonomically diverse lizard assemblage in Amazonian Ecuador. Canadian Journal of Zoology 74: 1313–1335.
  17. Vitt LJ, Zani PA, Avila-Pires TCS, Espósito MC (1998) Geographical ecology of the gymnophthalmid lizard Neusticurus ecpleopus in the Amazon rain forest. Canadian Journal of Zoology 76: 1671–1680. DOI: 10.1139/z98-114
  18. Valencia JH, Garzón-Tello K, Barragán-Paladines ME (2016) Serpientes venenosas del Ecuador: sistemática, taxonomía, historial natural, conservación, envenenamiento y aspectos antropológicos. Fundación Herpetológica Gustavo Orcés, Quito, 653 pp.
  19. Sherbrooke WC (1975) Reproductive cycle of a tropical teiid lizard, Neusticurus ecpleopus Cope, in Peru. Biotropica 7: 194–207. DOI: 10.2307/2989623
  20. Cisneros-Heredia DF, Calderón M (2019) Potamites ecpleopus. The IUCN Red List of threatened species. Available from: DOI: 10.2305/IUCN.UK.2019-2.RLTS.T44578695A44578704.en
  21. Zurita D (2015) Systematics of Amazonian lizards of the genus Potamites (Gymnophthalmidae). BSc thesis, Pontificia Universidad Católica del Ecuador, 74 pp.
  22. Cope ED (1875) Report on the reptiles brought by Professor James Orton from the middle and upper Amazon and western Peru. Journal of the Academy of Natural Sciences of Philadelphia 8: 159–183.

Appendix 1: Locality data used to create the distribution map of Potamites ecpleopus in Ecuador (Fig. 2). Go to the section on symbols and abbreviations for a list of acronyms used.

ColombiaCaquetáEl Manantial Nature and Community ReserveUAM-R-0403; SiB Colombia
ColombiaCaquetáLa ProfundaGutiérrez-Lamus et al. 2020
ColombiaCaquetáMonserrateGutiérrez-Lamus et al. 2020
ColombiaCaquetáParque Natural Municipal AndakíSINCHI-R 876; SiB Colombia
ColombiaCaquetáParque Natural Municipal Andakí, Los ÁngelesSINCHI-R 987; SiB Colombia
ColombiaPutumayoBetween Río Churuyaco and RumiyacoIAvH-R-4881; SiB Colombia
ColombiaPutumayoCascada Fin del MundoiNaturalist; photo examined
ColombiaPutumayoMocoaiNaturalist; photo examined
ColombiaPutumayoPuerto Asís, 5 km N of LACM 44670; SiB Colombia
ColombiaPutumayoVereda CampucanaPhoto by Angiie Tovar
EcuadorAzuaySopladora CampMZUA.Re.0223; examined
EcuadorMorona SantiagoArapicos, 1 km N ofRibeiro-Júnior & Amaral 2016
EcuadorMorona SantiagoCusuimeOrtega-Andrade 2010
EcuadorMorona SantiagoEl RosarioZurita 2015
EcuadorMorona SantiagoHuamboya, 7.4 km SE ofRibeiro-Júnior & Amaral 2016
EcuadorMorona SantiagoKapawi Jungle LodgeOrtega-Andrade 2010
EcuadorMorona SantiagoLas PeñasMZUA.Re.0210; examined
EcuadorMorona SantiagoMacumaiNaturalist; photo examined
EcuadorMorona SantiagoMacumaPhoto bank of the Reptiles of Ecuador book
EcuadorMorona SantiagoMiazalRibeiro-Júnior & Amaral 2016
EcuadorMorona SantiagoNapimiasTorres-Carvajal et al. 2016
EcuadorMorona SantiagoPaloraiNaturalist; photo examined
EcuadorMorona SantiagoParque Nacional SangayBrito & Almendariz 2013
EcuadorMorona SantiagoSantiago de MéndezOrtega-Andrade 2010
EcuadorMorona SantiagoSawastianOrtega-Andrade 2010
EcuadorMorona SantiagoSucúaRibeiro-Júnior & Amaral 2016
EcuadorMorona SantiagoVilla AshuaraOrtega-Andrade 2010
EcuadorMorona SantiagoWachirpas, 7 km E ofTorres-Carvajal et al. 2016
EcuadorMorona SantiagoWawaimi, 7.5 km E ofRibeiro-Júnior & Amaral 2016
EcuadorMorona SantiagoWisuiChaparro et al. 2011
EcuadorNapoCascada PuyupacchaiNaturalist; photo examined
EcuadorNapoCascada San RafaelRibeiro-Júnior & Amaral 2016
EcuadorNapoChontapunta, 4 km NE ofiNaturalist; photo examined
EcuadorNapoComuna 24 de MayoRibeiro-Júnior & Amaral 2016
EcuadorNapoHuaorani LodgePhoto bank of the Reptiles of Ecuador book
EcuadorNapoJondachi, 1.6 km W ofiNaturalist; photo examined
EcuadorNapoJondachi, 11 km E ofAvila-Pires 1995
EcuadorNapoJondachi, 13 km E ofAvila-Pires 1995
EcuadorNapoNarupa Biological ReserveMeza-Ramos et al. 2008
EcuadorNapoPuerto NapoMZUTI 1609; examined
EcuadorNapoRio Reventador, 2 km SW ofRibeiro-Júnior & Amaral 2016
EcuadorNapoSan Jose Viejo de SumacoUzzell 1996
EcuadorNapoSan Rafael, 1 km SW ofRibeiro-Júnior & Amaral 2016
EcuadorNapoSarayaku, CocodrilosPhoto bank of the Reptiles of Ecuador book
EcuadorNapoSendero abajo CocodrilosPhoto bank of the Reptiles of Ecuador book
EcuadorNapoTena, 6.7 km NW ofiNaturalist; photo examined
EcuadorNapoTena; 1 km E ofRibeiro-Júnior & Amaral 2016
EcuadorNapoWildsumaco LodgeCamper et al. 2021
EcuadorNapoWisdom Forest LodgeiNaturalist; photo examined
EcuadorNapoYachana ReserveWhitworth and Beirne 2011
EcuadorOrellanaBloque 31Libro PetroAmazonas
EcuadorOrellanaCocaRibeiro-Júnior & Amaral 2016
EcuadorOrellanaLoretoRibeiro-Júnior & Amaral 2016
EcuadorOrellanaMandaripanga LodgeReptiles of Ecuador book database
EcuadorOrellanaNWC Welcome CenterObservation by Alejandro Arteaga; this work
EcuadorOrellanaPompeya Sur–Iro roadChávez et al. 2021
EcuadorOrellanaPompeya Sur–Iro, km 22Torres-Carvajal et al. 2016
EcuadorOrellanaRío ArañaZurita 2015
EcuadorOrellanaRío BigalGarcía et al. 2021
EcuadorOrellanaRio CotopinoUzzell 1996
EcuadorOrellanaSan FranciscoZurita 2015
EcuadorOrellanaSan José de PayaminoMaynard et al. 2016
EcuadorOrellanaTiputini Biodiversity StationCisneros-Heredia 2006
EcuadorOrellanaYasuní National ParkCarvajal-Campos 2020
EcuadorOrellanaYasuní Research StationChávez et al. 2021
EcuadorOrellanaYuca SurPhoto by Paulina Romero
EcuadorOrellanaZancudocochaTorres-Carvajal et al. 2019
EcuadorPastaza10 de Agosto, 6.8 km N ofRibeiro-Júnior & Amaral 2016
EcuadorPastazaAndoasRibeiro-Júnior & Amaral 2016
EcuadorPastazaArutam Field StationSMF 90288; not examined
EcuadorPastazaBalsauraOrtega-Andrade 2010
EcuadorPastazaCabeceras del Río BobonazaAltamirano et al. 2014
EcuadorPastazaCachitambo, 8.4 km E ofRibeiro-Júnior & Amaral 2016
EcuadorPastazaComplejo Turístico los ChorritosiNaturalist; photo examined
EcuadorPastazaComunidad Copataza, on the Rio CopotazaAvila-Pires 1995
EcuadorPastazaComunidad Santa RosaRibeiro-Júnior & Amaral 2016
EcuadorPastazaConamboOrtega-Andrade 2010
EcuadorPastazaFátimaAvila-Pires 1995
EcuadorPastazaJuyuintzaOrtega-Andrade 2010
EcuadorPastazaKawymenoiNaturalist; photo examined
EcuadorPastazaKurintzaOrtega-Andrade 2010
EcuadorPastazaLorocachi, 3 km SE ofOrtega-Andrade 2010
EcuadorPastazaMeraUzzell 1996
EcuadorPastazaMera, 1.2 km N ofiNaturalist; photo examined
EcuadorPastazaMera, 7 km NW ofAvila-Pires 1995
EcuadorPastazaMera, 8 km NE ofiNaturalist; photo examined
EcuadorPastazaMontalvoRibeiro-Júnior & Amaral 2016
EcuadorPastazaMouth of Río CapahuariRibeiro-Júnior & Amaral 2016
EcuadorPastazaPeter Archer's placePhoto by Yatin Kalki
EcuadorPastazaPindoyakuOrtega-Andrade 2010
EcuadorPastazaPiwiriRibeiro-Júnior & Amaral 2016
EcuadorPastazaPuyoUSNM 203838; VertNet
EcuadorPastazaPuyo, 2 km SE ofiNaturalist; photo examined
EcuadorPastazaPuyo, 5 km N ofChávez et al. 2021
EcuadorPastazaRío Anzu Ecological ReserveReptiles of Ecuador book database
EcuadorPastazaRío CanelosRibeiro-Júnior & Amaral 2016
EcuadorPastazaRío Conambo and Río Pindayacu confluenceRibeiro-Júnior & Amaral 2016
EcuadorPastazaRío CopatazaRibeiro-Júnior & Amaral 2016
EcuadorPastazaRío OglánRibeiro-Júnior & Amaral 2016
EcuadorPastazaSantana Field StationSMF 90287; not examined
EcuadorPastazaSarayakuDoan & Castoe 2005
EcuadorPastazaSarayaku, on path to Río PalandayacuTorres-Carvajal et al. 2016
EcuadorPastazaShell, 2 km S ofRibeiro-Júnior & Amaral 2016
EcuadorPastazaShiripuno LodgeiNaturalist; photo examined
EcuadorPastazaSumak Kawsay In SituBentley et al. 2021
EcuadorPastazaTamandúa LodgePhoto by Jorge Flores
EcuadorPastazaTinajas del Río AnzuiNaturalist; photo examined
EcuadorPastazaTzarentzaZFMK 098717
EcuadorPastazaVeracruzRibeiro-Júnior & Amaral 2016
EcuadorSucumbíosAlma Ecuatoriana, 2 km W ofiNaturalist; photo examined
EcuadorSucumbíosCaiman LodgePhoto bank of the Reptiles of Ecuador book
EcuadorSucumbíosCascada de San Rafael, Cayambe-Coca Ecological ReserveChávez et al. 2021
EcuadorSucumbíosCascada San Rafael, 2 km NW ofiNaturalist; photo examined
EcuadorSucumbíosComunidad ZábaloCevallos Bustos 2010
EcuadorSucumbíosCuembi Triangle Protected ForestUTA 65509; not examined
EcuadorSucumbíosCuyabeno LodgeiNaturalist; photo examined
EcuadorSucumbíosDurenoiNaturalist; photo examined
EcuadorSucumbíosDureno, Río AguaricoValencia et al. 2016
EcuadorSucumbíosEl ReventadorAvila-Pires 1995
EcuadorSucumbíosGuacamayo EcolodgeiNaturalist; photo examined
EcuadorSucumbíosLa BarquillaYánez-Muñoz 2005
EcuadorSucumbíosLa BonitaZSFQ 2258
EcuadorSucumbíosLa Nueva TroncalKU 122192; VertNet
EcuadorSucumbíosLimoncocha Biological ReserveDoan & Castoe 2005
EcuadorSucumbíosLumbaquiRibeiro-Júnior & Amaral 2016
EcuadorSucumbíosLumbaqui, 5 km E ofPhoto bank of the Reptiles of Ecuador book
EcuadorSucumbíosNapo Wildlife CenterObservation by Alejandro Arteaga; this work
EcuadorSucumbíosNueva LojaKU 126810; VertNet
EcuadorSucumbíosNWC clay lickPhoto bank of the Reptiles of Ecuador book
EcuadorSucumbíosPalma RojaiNaturalist; photo examined
EcuadorSucumbíosRío Aguas BlancasYánez-Muñoz & Chimbo 2007
EcuadorSucumbíosRosa Florida, Cofan-Bermejo Reserve Buffer ZoneUTA 65511; not examined
EcuadorSucumbíosSacha LodgeiNaturalist; photo examined
EcuadorSucumbíosSan Pablo de KantesiyaMHNG 2237.076; collection database
EcuadorSucumbíosSani LodgeObservation by Alejandro Arteaga; this work
EcuadorSucumbíosSanta CeciliaBell et al. 2003
EcuadorSucumbíosSanta ElenaiNaturalist; photo examined
EcuadorTungurahuaCerro AbitaguaUzzell 1996
EcuadorTungurahuaRio Negro, 2.5 km NE of,MZUTI 4035; examined
EcuadorZamora ChinchipeAlto MachinazaAlmendáriz et al. 2014
EcuadorZamora ChinchipeColibríZurita 2015
EcuadorZamora ChinchipeValle dell QuimiBetancourt et al. 2018
PerúAmazonasAguaruna VillageBell et al. 2003
PerúAmazonasCaterpizaRibeiro-Júnior & Amaral 2016
PerúAmazonasChigkan EntseRibeiro-Júnior & Amaral 2016
PerúAmazonasCordillera del CóndorReynolds & Icochea 1997
PerúAmazonasHuambisa VillageBell et al. 2003
PerúAmazonasHuampamiBell et al. 2003
PerúAmazonasHuampami, 1 km SE ofMVZ 163186; VertNet
PerúAmazonasKusuBell et al. 2003
PerúAmazonasLa PozaBell et al. 2003
PerúAmazonasPaagatRibeiro-Júnior & Amaral 2016
PerúAmazonasQuebrada KampankisCatenazzi & Venegas 2012
PerúAmazonasQuebrada KaterpizaCatenazzi & Venegas 2012
PerúAmazonasQuebrada PastasilloBell et al. 2003
PerúAmazonasRio Kagka, headwaters ofRibeiro-Júnior & Amaral 2016
PerúAmazonasSan AntonioRibeiro-Júnior & Amaral 2016
PerúAmazonasShaimRibeiro-Júnior & Amaral 2016
PerúAmazonasShaimeRibeiro-Júnior & Amaral 2016
PerúAmazonasShiringaUSNM 568939; VertNet
PerúAmazonasSuaMVZ 163183; VertNet
PerúAmazonasTeniente PingloRibeiro-Júnior & Amaral 2016
PerúAmazonasTseasimRibeiro-Júnior & Amaral 2016
PerúAmazonasTuntanainMVZ 163194; VertNet
PerúAmazonasUpper Río ComainasRibeiro-Júnior & Amaral 2016
PerúAmazonasYutupisUSNM 583689; VertNet
PerúLoretoAguas NegrasYánez-Muñoz & Venegas 2008
PerúLoretoAndoasZurita 2015
PerúLoretoBellavista, 15 km S ofUSNM 565885; VertNet
PerúLoretoCampo AndoasValqui Schult 2015
PerúLoretoLoboyacu, 8 km N ofSERFOR
PerúLoretoPucacuro Reserved ZoneSERFOR
PerúLoretoPuerto Leguízamo, 19 km SW ofSERFOR
PerúLoretoQuebrada WeeCatenazzi & Venegas 2012
PerúLoretoRedondocochaYánez-Muñoz & Venegas 2008