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DOI10.47051/EMMQ9145 Published January 24, 2023. Updated February 8, 2024. Open access. Peer-reviewed.

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Common Stream-Lizard (Potamites ecpleopus)

Reptiles of Ecuador | Sauria | Gymnophthalmidae | Potamites ecpleopus

English common name: Common Stream-Lizard.

Spanish common name: Lagartija ribereña común.

Recognition: ♂♂ 17.6 cmMaximum distance from the snout to the tip of the tail. Snout–vent length=8.4 cm. ♀♀ 16.8 cmMaximum distance from the snout to the tip of the tail. Snout–vent length=7.1 cm..^1,2 Stream-Lizards of the genus Potamites are easily distinguishable from other leaf-litter lizards in their area of distribution by having heterogeneous dorsal scales and two longitudinal crests on the tail.³ The Common Stream-Lizard (P. ecpleopus) can be confused with its congener P. strangulatus and with species of the genus Gelanesaurus, from which it differs by having well-developed caudal ridges and enlarged tubercles around the neck, flanks, tail, and upper surface of the extremities, all of which give the lizard a crocodile-like appearance.^1–5 There is much geographic variation in P. ecpleopus.^4,6 In Ecuadorian populations, adult males have alternating black and white bars on the lips and edges of the chin, a dark throat coloration, and black ocelli with a cream center along the flanks. In females, the labial bars are dark brown instead of black, the throat is light gray, and there is a single ocellus on the insertion of the arm.^1,7 In adults, the dorsal coloration is tan to dull brown, whereas in juveniles the back is pinkish tan.¹

Figure 1: Individuals of Potamites ecpleopus: La Bonita, Sucumbíos province, Ecuador (); Palmarí Reserve, Amazonas state, Brazil (); Tzarentza, Pastaza province, Ecuador (); Río Bigai Reserve, Orellana province, Ecuador (). j=juvenile.

Natural history: In Ecuador, Potamites ecpleopus is considered a common lizard that occurs in high densities.^7,8 It inhabits old-growth rainforests both in flooded and terra-firme areas,^6–10 but also in disturbed areas such as pastures¹¹ and habitats affected by mining or logging.^2,8 These lizards are diurnal, semi-aquatic, and strongly associated with bodies of water.^4,9,12 They are good swimmers; in water, they propel themselves by means of sinuous movements of the body with help of the tail.^4,13 They also forage among leaf-litter, rocks, and logs along or around swampy areas, creeks, streams, and waterfalls,^7,14,15 usually partially submerged.^1,12 On occasion, these reptiles perch on vegetation up to almost 1 m above the ground or sit exposed on logs and rocks, basking in areas with filtered light.¹⁶ However, they prefer humid and shady areas within the forest and do not require direct sunlight to thermoregulate, being active during cloudy and rainy days, even in cold water.^1,9,17 At night, they roost on vegetation or among sticks and leaf-litter near streams.^8,13 They move slower than other co-occurring lizards and their body temperature is lower,^6,7 between 24–30°C, and directly influenced by the air or ground temperature.¹⁴

Individuals of Potamites ecpleopus return to the same point on consecutive days to feed.¹² Their diet is considered generalist and includes a wide variety of invertebrates ranging from earthworms to snails, but the most important prey items are crickets, ants, and insect larvae.^6,7 In smaller quantities, it also includes tadpoles¹⁷ and lizards such as Anolis trachyderma, Loxopholis parietalis, and even other P. ecpleopus.¹ There are records of snakes (Drymoluber dichrous,¹ Helicops angulatus,⁹ Oxyrhopus leucomelas,⁸ and Bothrocophias hyoprora¹⁸) preying upon individuals of P. ecpleopus. When threatened, these lizards run towards roots and leaf-litter, into holes in the banks,⁹ or directly to the water^4,7 submerging to the bottom debris.^1,6,14 If captured, they usually wriggle and bite fiercely, but can also shed the tail.^1,8 Potamites ecpleopus is an oviparous species with a continuous reproductive cycle.^9,19 The clutch size is two eggs measuring around 0.5 x 1.4 cm.² Communal nests of up to 5 eggs have been reported under large, rotten logs near bodies of water.⁴ Hatchlings measure 2.1–2.4 cm in SVL.⁹

Conservation: Least Concern Believed to be safe from extinction given current circumstances..²⁰ Potamites ecpleopus is listed in this category because it has a large distribution, occurs in protected areas, and has high population densities. Thus, the species is considered to be facing no major immediate extinction threats.²⁰ Although deforestation is affecting some populations, this is not considered a widespread threat to the species as a whole.²⁰ However, P. ecpleopus has been shown to be a species complex²¹ and some populations are under greater threat from habitat loss.

Distribution: Potamites ecpleopus is widely distributed throughout the Amazon basin of Bolivia, Brazil, Colombia, Ecuador (Fig. 2), and Perú.² It also occurs along the adjacent Andean foothills.

Figure 2: Distribution of Potamites ecpleopus in Ecuador. See Appendix 1 for a complete list of the presence localities included in the map.

Etymology: The generic name Potamites, which is a Greek noun meaning “water finder,” refers to the semi-aquatic habits of lizards in this genus.³ The specific epithet ecpleopus refers to the similarity between the head of this species and those of the tribe Ecpleopini.²²

See it in the wild: Common Stream-Lizards can be observed with almost complete certainty in forested streams throughout the species’ area of distribution in Ecuador. These reptiles are particularly common in Yasuní National Park, Sumaco National Park, and Cuyabeno Wildlife Reserve. Individuals can be found by removing leaf-litter or logs along streams or flooded areas within the forest

Authors: Amanda Quezada^aAffiliation: Tropical Herping (TH), Quito, Ecuador.^,bAffiliation: Laboratorio de Herpetología, Universidad del Azuay, Cuenca, Ecuador. and Alejandro Arteaga^cAffiliation: Khamai Foundation, Quito, Ecuador.

Academic reviewer: Jeffrey D Camper^dAffiliation: Department of Biology, Francis Marion University, Florence, USA.

Photographers: Jose Vieira^aAffiliation: Tropical Herping (TH), Quito, Ecuador.^,eAffiliation: ExSitu, Quito, Ecuador. and Sebastián Di Doménico^fAffiliation: Keeping Nature, Bogotá, Colombia.

How to cite? Quezada A, Arteaga A (2024) Common Stream-Lizard (Potamites ecpleopus). In: Arteaga A, Bustamante L, Vieira J (Eds) Reptiles of Ecuador: Life in the middle of the world. Available from: www.reptilesofecuador.com. DOI: 10.47051/EMMQ9145

Literature cited:

1. Duellman WE (1978) The biology of an equatorial herpetofauna in Amazonian Ecuador. Publications of the Museum of Natural History, University of Kansas 65: 1–352.
2. Avila-Pires TCS (1995) Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen 299: 1–706.
3. Doan TM, Castoe TA (2005) Phylogenetic taxonomy of the Cercosaurini (Squamata: Gymnophthalmidae), with new genera for species of Neusticurus and Proctoporus. Zoological Journal of the Linnean Society 143: 405–416. DOI: 10.1111/j.1096-3642.2005.00145.x
4. Uzzell TM (1966) Teid Lizards of the genus Neusticurus (Reptilia, Sauria). Bulletin of the American Museum of Natural History 132: 279–327.
5. Altamirano-Benavides M, Zaher H, Lobo L, Grazziotin FG, Sales Nunes PM, Rodrigues MT (2013) A new species of lizard genus Potamites from Ecuador (Squamata, Gymnophthalmidae). Zootaxa 3717: 345–358. DOI: 10.11646/zootaxa.3717.3.4
6. Vitt LJ, Avila-Pires TCS (1998) Ecology of two sympatric species of Neusticurus (Sauria: Gymnophthalmidae) in the Western Amazon of Brazil. Copeia 1998: 570–582. DOI: 10.2307/1447787
7. Vitt LJ, De la Torre S (1996) A research guide to the lizards of Cuyabeno. Pontificia Universidad Católica del Ecuador, Quito, 165 pp.
8. Field notes, Reptiles of Ecuador book project.
9. Dixon JR, Soini P (1986) The reptiles of the upper Amazon Basin, Iquitos region, Peru. Milwaukee Public Museum, Milwaukee, 154 pp.
10. Catenazzi A, Lehr E, von May R (2013) The amphibians and reptiles of Manu National Park and its buffer zone, Amazon basin and eastern slopes of the Andes, Peru. Biota Neotropica 13: 269–283. DOI: 10.1590/S1676-06032013000400024
11. Camper JD, Torres-Carvajal O, Ron SR, Nilsson J, Arteaga A, Knowles TW, Arbogast BS (2021) Amphibians and reptiles of Wildsumaco Wildlife Sanctuary, Napo Province, Ecuador. Check List 17: 729–751.
12. Rocha CFD (1991) Habitat utilization and feeding habits of Neusticurus ecpleopus in a Brazilian tropical rainforest. Herpetological Review 22: 40–42.
13. Whitworth A, Beirne C (2011) Reptiles of the Yachana Reserve. Global Vision International, Exeter, 130 pp.
14. Fitch HS (1968) Temperature and behavior of some equatorial lizards. Herpetologica 24: 35–38.
15. da Fonseca WL, da Silva JD, Abegg AD, da Rosa CM, Bernarde PS (2019) Herpetofauna of Porto Walter and surrounding areas, southwest Amazonia, Brazil. Herpetology Notes 12: 91–107.
16. Vitt LJ, Zani PA (1996) Organization of a taxonomically diverse lizard assemblage in Amazonian Ecuador. Canadian Journal of Zoology 74: 1313–1335.
17. Vitt LJ, Zani PA, Avila-Pires TCS, Espósito MC (1998) Geographical ecology of the gymnophthalmid lizard Neusticurus ecpleopus in the Amazon rain forest. Canadian Journal of Zoology 76: 1671–1680. DOI: 10.1139/z98-114
18. Valencia JH, Garzón-Tello K, Barragán-Paladines ME (2016) Serpientes venenosas del Ecuador: sistemática, taxonomía, historial natural, conservación, envenenamiento y aspectos antropológicos. Fundación Herpetológica Gustavo Orcés, Quito, 653 pp.
19. Sherbrooke WC (1975) Reproductive cycle of a tropical teiid lizard, Neusticurus ecpleopus Cope, in Peru. Biotropica 7: 194–207. DOI: 10.2307/2989623
20. Cisneros-Heredia DF, Calderón M (2019) Potamites ecpleopus. The IUCN Red List of threatened species. Available from: www.iucnredlist.org. DOI: 10.2305/IUCN.UK.2019-2.RLTS.T44578695A44578704.en
21. Zurita D (2015) Systematics of Amazonian lizards of the genus Potamites (Gymnophthalmidae). BSc thesis, Pontificia Universidad Católica del Ecuador, 74 pp.
22. Cope ED (1875) Report on the reptiles brought by Professor James Orton from the middle and upper Amazon and western Peru. Journal of the Academy of Natural Sciences of Philadelphia 8: 159–183.

Appendix 1: Locality data used to create the distribution map of Potamites ecpleopus in Ecuador (Fig. 2). Go to the section on symbols and abbreviations for a list of acronyms used.

Country	Province	Locality	Source
Colombia	Caquetá	El Manantial Nature and Community Reserve	UAM-R-0403; SiB Colombia
Colombia	Caquetá	La Profunda	Gutiérrez-Lamus et al. 2020
Colombia	Caquetá	Monserrate	Gutiérrez-Lamus et al. 2020
Colombia	Caquetá	Parque Natural Municipal Andakí	SINCHI-R 876; SiB Colombia
Colombia	Caquetá	Parque Natural Municipal Andakí, Los Ángeles	SINCHI-R 987; SiB Colombia
Colombia	Putumayo	Between Río Churuyaco and Rumiyaco	IAvH-R-4881; SiB Colombia
Colombia	Putumayo	Cascada Fin del Mundo	iNaturalist; photo examined
Colombia	Putumayo	Mocoa	iNaturalist; photo examined
Colombia	Putumayo	Puerto Asís, 5 km N of	LACM 44670; SiB Colombia
Colombia	Putumayo	Vereda Campucana	Photo by Angiie Tovar
Ecuador	Azuay	Sopladora Camp	MZUA.Re.0223; examined
Ecuador	Morona Santiago	Arapicos, 1 km N of	Ribeiro-Júnior & Amaral 2016
Ecuador	Morona Santiago	Cusuime	Ortega-Andrade 2010
Ecuador	Morona Santiago	El Rosario	Zurita 2015
Ecuador	Morona Santiago	Huamboya, 7.4 km SE of	Ribeiro-Júnior & Amaral 2016
Ecuador	Morona Santiago	Kapawi Jungle Lodge	Ortega-Andrade 2010
Ecuador	Morona Santiago	Las Peñas	MZUA.Re.0210; examined
Ecuador	Morona Santiago	Macuma	iNaturalist; photo examined
Ecuador	Morona Santiago	Macuma	Photo bank of the Reptiles of Ecuador book
Ecuador	Morona Santiago	Miazal	Ribeiro-Júnior & Amaral 2016
Ecuador	Morona Santiago	Napimias	Torres-Carvajal et al. 2016
Ecuador	Morona Santiago	Palora	iNaturalist; photo examined
Ecuador	Morona Santiago	Parque Nacional Sangay	Brito & Almendariz 2013
Ecuador	Morona Santiago	Santiago de Méndez	Ortega-Andrade 2010
Ecuador	Morona Santiago	Sawastian	Ortega-Andrade 2010
Ecuador	Morona Santiago	Sucúa	Ribeiro-Júnior & Amaral 2016
Ecuador	Morona Santiago	Villa Ashuara	Ortega-Andrade 2010
Ecuador	Morona Santiago	Wachirpas, 7 km E of	Torres-Carvajal et al. 2016
Ecuador	Morona Santiago	Wawaimi, 7.5 km E of	Ribeiro-Júnior & Amaral 2016
Ecuador	Morona Santiago	Wisui	Chaparro et al. 2011
Ecuador	Napo	Cascada Puyupaccha	iNaturalist; photo examined
Ecuador	Napo	Cascada San Rafael	Ribeiro-Júnior & Amaral 2016
Ecuador	Napo	Chontapunta, 4 km NE of	iNaturalist; photo examined
Ecuador	Napo	Comuna 24 de Mayo	Ribeiro-Júnior & Amaral 2016
Ecuador	Napo	Huaorani Lodge	Photo bank of the Reptiles of Ecuador book
Ecuador	Napo	Jondachi, 1.6 km W of	iNaturalist; photo examined
Ecuador	Napo	Jondachi, 11 km E of	Avila-Pires 1995
Ecuador	Napo	Jondachi, 13 km E of	Avila-Pires 1995
Ecuador	Napo	Narupa Biological Reserve	Meza-Ramos et al. 2008
Ecuador	Napo	Puerto Napo	MZUTI 1609; examined
Ecuador	Napo	Rio Reventador, 2 km SW of	Ribeiro-Júnior & Amaral 2016
Ecuador	Napo	San Jose Viejo de Sumaco	Uzzell 1996
Ecuador	Napo	San Rafael, 1 km SW of	Ribeiro-Júnior & Amaral 2016
Ecuador	Napo	Sarayaku, Cocodrilos	Photo bank of the Reptiles of Ecuador book
Ecuador	Napo	Sendero abajo Cocodrilos	Photo bank of the Reptiles of Ecuador book
Ecuador	Napo	Tena, 6.7 km NW of	iNaturalist; photo examined
Ecuador	Napo	Tena; 1 km E of	Ribeiro-Júnior & Amaral 2016
Ecuador	Napo	Wildsumaco Lodge	Camper et al. 2021
Ecuador	Napo	Wisdom Forest Lodge	iNaturalist; photo examined
Ecuador	Napo	Yachana Reserve	Whitworth and Beirne 2011
Ecuador	Orellana	Bloque 31	Libro PetroAmazonas
Ecuador	Orellana	Coca	Ribeiro-Júnior & Amaral 2016
Ecuador	Orellana	Loreto	Ribeiro-Júnior & Amaral 2016
Ecuador	Orellana	Mandaripanga Lodge	Reptiles of Ecuador book database
Ecuador	Orellana	NWC Welcome Center	Observation by Alejandro Arteaga; this work
Ecuador	Orellana	Pompeya Sur–Iro road	Chávez et al. 2021
Ecuador	Orellana	Pompeya Sur–Iro, km 22	Torres-Carvajal et al. 2016
Ecuador	Orellana	Río Araña	Zurita 2015
Ecuador	Orellana	Río Bigal	García et al. 2021
Ecuador	Orellana	Rio Cotopino	Uzzell 1996
Ecuador	Orellana	San Francisco	Zurita 2015
Ecuador	Orellana	San José de Payamino	Maynard et al. 2016
Ecuador	Orellana	Tiputini Biodiversity Station	Cisneros-Heredia 2006
Ecuador	Orellana	Yasuní National Park	Carvajal-Campos 2020
Ecuador	Orellana	Yasuní Research Station	Chávez et al. 2021
Ecuador	Orellana	Yuca Sur	Photo by Paulina Romero
Ecuador	Orellana	Zancudococha	Torres-Carvajal et al. 2019
Ecuador	Pastaza	10 de Agosto, 6.8 km N of	Ribeiro-Júnior & Amaral 2016
Ecuador	Pastaza	Andoas	Ribeiro-Júnior & Amaral 2016
Ecuador	Pastaza	Arutam Field Station	SMF 90288; not examined
Ecuador	Pastaza	Balsaura	Ortega-Andrade 2010
Ecuador	Pastaza	Cabeceras del Río Bobonaza	Altamirano et al. 2014
Ecuador	Pastaza	Cachitambo, 8.4 km E of	Ribeiro-Júnior & Amaral 2016
Ecuador	Pastaza	Complejo Turístico los Chorritos	iNaturalist; photo examined
Ecuador	Pastaza	Comunidad Copataza, on the Rio Copotaza	Avila-Pires 1995
Ecuador	Pastaza	Comunidad Santa Rosa	Ribeiro-Júnior & Amaral 2016
Ecuador	Pastaza	Conambo	Ortega-Andrade 2010
Ecuador	Pastaza	Fátima	Avila-Pires 1995
Ecuador	Pastaza	Juyuintza	Ortega-Andrade 2010
Ecuador	Pastaza	Kawymeno	iNaturalist; photo examined
Ecuador	Pastaza	Kurintza	Ortega-Andrade 2010
Ecuador	Pastaza	Lorocachi, 3 km SE of	Ortega-Andrade 2010
Ecuador	Pastaza	Mera	Uzzell 1996
Ecuador	Pastaza	Mera, 1.2 km N of	iNaturalist; photo examined
Ecuador	Pastaza	Mera, 7 km NW of	Avila-Pires 1995
Ecuador	Pastaza	Mera, 8 km NE of	iNaturalist; photo examined
Ecuador	Pastaza	Montalvo	Ribeiro-Júnior & Amaral 2016
Ecuador	Pastaza	Mouth of Río Capahuari	Ribeiro-Júnior & Amaral 2016
Ecuador	Pastaza	Peter Archer's place	Photo by Yatin Kalki
Ecuador	Pastaza	Pindoyaku	Ortega-Andrade 2010
Ecuador	Pastaza	Piwiri	Ribeiro-Júnior & Amaral 2016
Ecuador	Pastaza	Puyo	USNM 203838; VertNet
Ecuador	Pastaza	Puyo, 2 km SE of	iNaturalist; photo examined
Ecuador	Pastaza	Puyo, 5 km N of	Chávez et al. 2021
Ecuador	Pastaza	Río Anzu Ecological Reserve	Reptiles of Ecuador book database
Ecuador	Pastaza	Río Canelos	Ribeiro-Júnior & Amaral 2016
Ecuador	Pastaza	Río Conambo and Río Pindayacu confluence	Ribeiro-Júnior & Amaral 2016
Ecuador	Pastaza	Río Copataza	Ribeiro-Júnior & Amaral 2016
Ecuador	Pastaza	Río Oglán	Ribeiro-Júnior & Amaral 2016
Ecuador	Pastaza	Santana Field Station	SMF 90287; not examined
Ecuador	Pastaza	Sarayaku	Doan & Castoe 2005
Ecuador	Pastaza	Sarayaku, on path to Río Palandayacu	Torres-Carvajal et al. 2016
Ecuador	Pastaza	Shell, 2 km S of	Ribeiro-Júnior & Amaral 2016
Ecuador	Pastaza	Shiripuno Lodge	iNaturalist; photo examined
Ecuador	Pastaza	Sumak Kawsay In Situ	Bentley et al. 2021
Ecuador	Pastaza	Tamandúa Lodge	Photo by Jorge Flores
Ecuador	Pastaza	Tinajas del Río Anzu	iNaturalist; photo examined
Ecuador	Pastaza	Tzarentza	ZFMK 098717
Ecuador	Pastaza	Veracruz	Ribeiro-Júnior & Amaral 2016
Ecuador	Sucumbíos	Alma Ecuatoriana, 2 km W of	iNaturalist; photo examined
Ecuador	Sucumbíos	Caiman Lodge	Photo bank of the Reptiles of Ecuador book
Ecuador	Sucumbíos	Cascada de San Rafael, Cayambe-Coca Ecological Reserve	Chávez et al. 2021
Ecuador	Sucumbíos	Cascada San Rafael, 2 km NW of	iNaturalist; photo examined
Ecuador	Sucumbíos	Comunidad Zábalo	Cevallos Bustos 2010
Ecuador	Sucumbíos	Cuembi Triangle Protected Forest	UTA 65509; not examined
Ecuador	Sucumbíos	Cuyabeno Lodge	iNaturalist; photo examined
Ecuador	Sucumbíos	Dureno	iNaturalist; photo examined
Ecuador	Sucumbíos	Dureno, Río Aguarico	Valencia et al. 2016
Ecuador	Sucumbíos	El Reventador	Avila-Pires 1995
Ecuador	Sucumbíos	Guacamayo Ecolodge	iNaturalist; photo examined
Ecuador	Sucumbíos	La Barquilla	Yánez-Muñoz 2005
Ecuador	Sucumbíos	La Bonita	ZSFQ 2258
Ecuador	Sucumbíos	La Nueva Troncal	KU 122192; VertNet
Ecuador	Sucumbíos	Limoncocha Biological Reserve	Doan & Castoe 2005
Ecuador	Sucumbíos	Lumbaqui	Ribeiro-Júnior & Amaral 2016
Ecuador	Sucumbíos	Lumbaqui, 5 km E of	Photo bank of the Reptiles of Ecuador book
Ecuador	Sucumbíos	Napo Wildlife Center	Observation by Alejandro Arteaga; this work
Ecuador	Sucumbíos	Nueva Loja	KU 126810; VertNet
Ecuador	Sucumbíos	NWC clay lick	Photo bank of the Reptiles of Ecuador book
Ecuador	Sucumbíos	Palma Roja	iNaturalist; photo examined
Ecuador	Sucumbíos	Río Aguas Blancas	Yánez-Muñoz & Chimbo 2007
Ecuador	Sucumbíos	Rosa Florida, Cofan-Bermejo Reserve Buffer Zone	UTA 65511; not examined
Ecuador	Sucumbíos	Sacha Lodge	iNaturalist; photo examined
Ecuador	Sucumbíos	San Pablo de Kantesiya	MHNG 2237.076; collection database
Ecuador	Sucumbíos	Sani Lodge	Observation by Alejandro Arteaga; this work
Ecuador	Sucumbíos	Santa Cecilia	Bell et al. 2003
Ecuador	Sucumbíos	Santa Elena	iNaturalist; photo examined
Ecuador	Tungurahua	Cerro Abitagua	Uzzell 1996
Ecuador	Tungurahua	Rio Negro, 2.5 km NE of,	MZUTI 4035; examined
Ecuador	Zamora Chinchipe	Alto Machinaza	Almendáriz et al. 2014
Ecuador	Zamora Chinchipe	Colibrí	Zurita 2015
Ecuador	Zamora Chinchipe	Valle dell Quimi	Betancourt et al. 2018
Perú	Amazonas	Aguaruna Village	Bell et al. 2003
Perú	Amazonas	Caterpiza	Ribeiro-Júnior & Amaral 2016
Perú	Amazonas	Chigkan Entse	Ribeiro-Júnior & Amaral 2016
Perú	Amazonas	Cordillera del Cóndor	Reynolds & Icochea 1997
Perú	Amazonas	Huambisa Village	Bell et al. 2003
Perú	Amazonas	Huampami	Bell et al. 2003
Perú	Amazonas	Huampami, 1 km SE of	MVZ 163186; VertNet
Perú	Amazonas	Kusu	Bell et al. 2003
Perú	Amazonas	La Poza	Bell et al. 2003
Perú	Amazonas	Paagat	Ribeiro-Júnior & Amaral 2016
Perú	Amazonas	Quebrada Kampankis	Catenazzi & Venegas 2012
Perú	Amazonas	Quebrada Katerpiza	Catenazzi & Venegas 2012
Perú	Amazonas	Quebrada Pastasillo	Bell et al. 2003
Perú	Amazonas	Rio Kagka, headwaters of	Ribeiro-Júnior & Amaral 2016
Perú	Amazonas	San Antonio	Ribeiro-Júnior & Amaral 2016
Perú	Amazonas	Shaim	Ribeiro-Júnior & Amaral 2016
Perú	Amazonas	Shaime	Ribeiro-Júnior & Amaral 2016
Perú	Amazonas	Shiringa	USNM 568939; VertNet
Perú	Amazonas	Sua	MVZ 163183; VertNet
Perú	Amazonas	Teniente Pinglo	Ribeiro-Júnior & Amaral 2016
Perú	Amazonas	Tseasim	Ribeiro-Júnior & Amaral 2016
Perú	Amazonas	Tuntanain	MVZ 163194; VertNet
Perú	Amazonas	Upper Río Comainas	Ribeiro-Júnior & Amaral 2016
Perú	Amazonas	Yutupis	USNM 583689; VertNet
Perú	Loreto	Aguas Negras	Yánez-Muñoz & Venegas 2008
Perú	Loreto	Andoas	Zurita 2015
Perú	Loreto	Bellavista, 15 km S of	USNM 565885; VertNet
Perú	Loreto	Campo Andoas	Valqui Schult 2015
Perú	Loreto	Loboyacu, 8 km N of	SERFOR
Perú	Loreto	Pucacuro Reserved Zone	SERFOR
Perú	Loreto	Puerto Leguízamo, 19 km SW of	SERFOR
Perú	Loreto	Quebrada Wee	Catenazzi & Venegas 2012
Perú	Loreto	Redondococha	Yánez-Muñoz & Venegas 2008

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