Published June 18, 2021. Updated November 7, 2023. Open access. | Purchase book ❯ |
Yellow-tongued Anole (Anolis scypheus)
Reptiles of Ecuador | Sauria | Anolidae | Anolis scypheus
English common names: Yellow-tongued Anole, Dim Anole.
Spanish common names: Anolis de lengua amarilla, anolis cabeza de platillo, anolis sombrío.
Recognition: ♂♂ 26.6 cmMaximum distance from the snout to the tip of the tail. Snout–vent length=8.4 cm. ♀♀ 25.6 cmMaximum distance from the snout to the tip of the tail. Snout–vent length=8.6 cm..1 Anoles are easily distinguishable from other lizards by their diurnal habits, extensible dewlap in males, expanded digital pads, and granular scales on the dorsum and belly.2 Yellow-tongued Anoles (Anolis scypheus) can be recognized by being moderately robust, having a short snout, and a dewlap that is blue medially and red peripherally.1,3,4 The coloration resembles that of a dry leaf, with cream caudally directed chevrons and streaks on a dark brown dorsum.5,6 Females differ from males by having a smaller dewlap, and in some cases, a pale vertebral stripe.1,4 Anolis scypheus resembles A. bombiceps in size and dorsal coloration, but the latter has an entirely blue dewlap.5
Natural history: Anolis scypheus is a diurnal and primarily terrestrial lizard adapted to life on the leaf-litter and understory vegetation.4 The species is compartively abundant in old-growth to heavily disturbed evergreen lowland forests, including seasonally flooded as well as terra-firme areas. These anoles forage mostly at ground level7 or on tree trunks and shrubs less than 2.5 m above the ground.3,8 Their activity occurs throughout the day and mostly under shade,4 but individuals have also been seen basking on logs or buttresses.1 At night, they sleep on leaves, sticks, and stems less than 1.5 m above the ground.4,9 Yellow-tongued Anoles are ambush predators that attack mobile insects that pass nearby.1,4 They feed mostly on invertebrates such as grasshoppers, katydids, roaches, insect larvae, spiders, and isopods, but may as well include lizards (such as A. trachyderma and Loxopholis parietalis) in their diet.1,4 Yellow-tongued Anoles escape predators by remaining perfectly still; which, combined with their dry leaf camouflage, makes them hard to detect.4,9 When disturbed, they may remain motionless or run for a few meters and disappear in the shade.10 If on a trunk, they move to the opposite side or run up along the trunk, sometimes up to 9 m above the ground.4,9 There is an unpublished record of a snake (Imantodes lentiferus) preying upon an individual of A. scypheus.9 The breeding season appears to take place year-round.1 Gravid females contain 1–2 eggs,1 but the clutch consists of one egg at a time.4
Conservation: Least Concern Believed to be safe from extinction given current circumstances..11,12 Anolis scypheus is listed in this category given its wide distribution, presence in major protected areas, lack of widespread threats, and presumed large stable populations.11 The species occurs over areas that retain the majority of their vegetation cover. In Brazil, 52% of the occurrence area of the species is inside protected areas, and about 99.8% of its distribution still holds continuous forest cover.13 In Ecuador, an estimated 89% of the habitat of the species is still forested.14
Distribution: Anolis scypheus is widely distributed throughout western Amazonia in Brazil, Colombia, Ecuador (Fig. 2), and Perú.15 It also ranges along the Andean-Orinoquian foothills of Venezuela and Colombia.7
Etymology: The generic name Anolis is thought to have originated from Cariban languages, specifically from the word anoli, which is the name Arawak peoples may have used to refer to this group of lizards.16 The specific epithet scypheus may be derived from the Greek word skyphion (=dim),17 in reference to the dorsal coloration,18 or from the Latin scyphus (=cup),19 in reference to the shape of the head.18
See it in the wild: Yellow-tongued Anoles can be found in forested areas throughout their distribution in Ecuador, at a rate of about once every two days. Prime locations to find these lizards in Ecuador include the Yasuní Scientific Station, Limoncocha Biological Reserve, Shiripuno Lodge, Huella Verde Lodge, and Jatun Sacha Biological Reserve. While it is possible to spot individuals actively moving about on the forest floor during the day, they are more readily observed and approached at night when they are sleeping on twigs and leaves near the ground.
Acknowledgments: This account was published with the support of Secretaría Nacional de Educación Superior Ciencia y Tecnología (programa INEDITA; project: Respuestas a la crisis de biodiversidad: la descripción de especies como herramienta de conservación; No 00110378), Programa de las Naciones Unidas (PNUD), and Universidad San Francisco de Quito (USFQ).
Special thanks to David Turell for symbolically adopting the Yellow-tongued Anole and helping bring the Reptiles of Ecuador book project to life.
Click here to adopt a species.
Author: Alejandro ArteagaaAffiliation: Khamai Foundation, Quito, Ecuador.
Photographers: Alejandro Arteaga,dAffiliation: Tropical Herping (TH), Quito, Ecuador. Jose Vieira,aAffiliation: Tropical Herping (TH), Quito, Ecuador.,bAffiliation: ExSitu, Quito, Ecuador. and Sebastián Di DoménicocAffiliation: Keeping Nature, Bogotá, Colombia.
How to cite? Arteaga A (2021) Yellow-tongued Anole (Anolis scypheus). In: Arteaga A, Bustamante L, Vieira J (Eds) Reptiles of Ecuador: Life in the middle of the world. Available from: www.reptilesofecuador.com. DOI: 10.47051/YVMA4242
Literature cited:
- Duellman WE (1978) The biology of an equatorial herpetofauna in Amazonian Ecuador. Publications of the Museum of Natural History, University of Kansas 65: 1–352.
- Peters JA, Donoso-Barros R (1970) Catalogue of the Neotropical Squamata: part II, lizards and amphisbaenians. Bulletin of the United States National Museum, Washington, D.C., 293 pp.
- Dixon JR, Soini P (1986) The reptiles of the upper Amazon Basin, Iquitos region, Peru. Milwaukee Public Museum, Milwaukee, 154 pp.
- Vitt LJ, De la Torre S (1996) A research guide to the lizards of Cuyabeno. Pontificia Universidad Católica del Ecuador, Quito, 165 pp.
- D’Angiolella AB, Gamble T, Avila-Pires TCS, Colli GR, Noonan BP, Vitt LJ (2011) Anolis chrysolepis Duméril and Bibron, 1837 (Squamata: Iguanidae), revisited: molecular phylogeny and taxonomy of the Anolis chrysolepis species group. Bulletin of the Museum of Comparative Zoology 160: 35–63. DOI: 10.3099/0027-4100-160.2.35
- Avila-Pires TCS (1995) Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen 299: 1–706.
- Rojas-Runjaic FJ, Quihua D, Castellanos M, La Marca E (2018) Anolis scypheus Cope, 1864 (Squamata, Dactyloidae) en Venezuela: confirmación de presencia y nuevos registros. Memoria de la Fundación La Salle de Ciencias Naturales 76: 93–101.
- Vitt LJ, Zani PA (1996) Organization of a taxonomically diverse lizard assemblage in Amazonian Ecuador. Canadian Journal of Zoology 74: 1313–1335.
- Field notes, Reptiles of Ecuador book project.
- Fitch HS (1968) Temperature and behavior of some equatorial lizards. Herpetologica 24: 35–38.
- Caicedo JR, Calderón M, Ines Hladki A, Ramírez Pinilla M, Renjifo J, Rivas G, Urbina N, Perez P, Avila-Pires TCS, Cisneros-Heredia DF (2020) Anolis scypheus. The IUCN Red List of threatened species. Available from: www.iucnredlist.org. DOI: 10.2305/IUCN.UK.2020-3.RLTS.T44577756A44577761.en
- Reyes-Puig C (2015) Un método integrativo para evaluar el estado de conservación de las especies y su aplicación a los reptiles del Ecuador. MSc thesis, Pontificia Universidad Católica del Ecuador, 73 pp.
- Ribeiro-Júnior MA, Amaral S (2016) Diversity, distribution, and conservation of lizards (Reptilia: Squamata) in the Brazilian Amazonia. Neotropical Biodiversity 2: 195–421. DOI: 10.1080/23766808.2016.1236769
- MAE (2012) Línea base de deforestación del Ecuador continental. Ministerio del Ambiente del Ecuador, Quito, 30 pp.
- Ribeiro-Junior MA (2015) Catalogue of distribution of lizards (Reptilia: Squamata) from the Brazilian Amazonia. I. Dactyloidae, Hoplocercidae, Iguanidae, Leiosauridae, Polychrotidae, Tropiduridae. Zootaxa 3983: 001–110. DOI: 10.11646/zootaxa.3983.1.1
- Allsopp R (1996) Dictionary of Caribbean English Usage. Oxford University Press, Oxford, 776 pp.
- Brown RW (1956) Composition of scientific words. Smithsonian Books, Washington D.C., 882 pp.
- Cope ED (1864) Contributions to the herpetology of tropical America. Proceedings of the Academy of Natural Sciences of Philadelphia 16: 166–181.
- Mir J (1982) Diccionario ilustrado Latín. Barcelona, 557 pp.
Appendix 1: Locality data used to create the distribution map of Anolis scypheus in Ecuador (Fig. 2). Go to the section on symbols and abbreviations for a list of acronyms used.
Country | Province | Locality | Source |
Colombia | Caquetá | Florencia | Photo by Camilo Yasnó |
Colombia | Caquetá | Monserrate | Gutiérrez-Lamus et al. 2020 |
Colombia | Putumayo | Orito | Ribeiro-Júnior 2015 |
Colombia | Putumayo | Río Picudo Chico | IAvH 392 |
Colombia | Putumayo | Río Putumayo | FMNH 165808 |
Colombia | Putumayo | Vereda Alpes Orientales | IAvH 330 |
Ecuador | Morona Santiago | Centro Numpaim | Torres-Carvajal et al. 2019 |
Ecuador | Morona Santiago | Miazal | Ribeiro-Júnior 2015 |
Ecuador | Morona Santiago | Río Cusuime | Ribeiro-Júnior 2015 |
Ecuador | Morona Santiago | Río Llushin | Ribeiro-Júnior 2015 |
Ecuador | Morona Santiago | Río Upano | Ribeiro-Júnior 2015 |
Ecuador | Morona Santiago | Sawastian | Ortega-Andrade 2010 |
Ecuador | Morona Santiago | Shuin Mamus | iNaturalist |
Ecuador | Morona Santiago | Soldado Monge | Photo by Freddy Velásquez |
Ecuador | Morona Santiago | Sucúa, 2 miles E of | Ribeiro-Júnior 2015 |
Ecuador | Morona Santiago | Unión del Río Paute y Río Negro | Torres-Carvajal et al. 2019 |
Ecuador | Napo | Asociación Tsatsayaku | iNaturalist |
Ecuador | Napo | Gareno Lodge | This work |
Ecuador | Napo | Huaorani Lodge | This work |
Ecuador | Napo | Jatun Sacha Biological Station | This work |
Ecuador | Napo | La Cruz Blanca | D’Angiolella et al. 2011 |
Ecuador | Napo | Liana Lodge | This work |
Ecuador | Napo | Río Misahuallí | Ribeiro-Júnior 2015 |
Ecuador | Napo | Sacha Lodge | This work |
Ecuador | Napo | Yachana Reserve | Whitworth & Beirne 2011 |
Ecuador | Orellana | Bloque 31 | Libro PetroAmazonas |
Ecuador | Orellana | Boya 2 | MECN 3380 |
Ecuador | Orellana | Dayuma, 22.6 km E of | MECN 1934 |
Ecuador | Orellana | El Coca | MHNG 2591.021 |
Ecuador | Orellana | Estación Pindo | iNaturalist |
Ecuador | Orellana | Hacienda Primavera | D’Angiolella et al. 2011 |
Ecuador | Orellana | Joya de los Sachas | MECN 4989 |
Ecuador | Orellana | Laguna Taracoa | D’Angiolella et al. 2011 |
Ecuador | Orellana | Mandaripanga Lodge | iNaturalist |
Ecuador | Orellana | Napo Wildlife Center | iNaturalist |
Ecuador | Orellana | Obe Occidental | MECN 3383 |
Ecuador | Orellana | Paushiyacu | Kingsbury et al. 2008 |
Ecuador | Orellana | Pozo Amo 2 | Ribeiro-Júnior 2015 |
Ecuador | Orellana | Río Bigal Biological Reserve | Thierry García |
Ecuador | Orellana | Río Yasuní | This work |
Ecuador | Orellana | San Jose de Mote | Ribeiro-Júnior 2015 |
Ecuador | Orellana | Shiripuno Lodge | iNaturalist |
Ecuador | Orellana | Tiguino | MECN 383 |
Ecuador | Orellana | Tiputini Biodiversity Station | iNaturalist |
Ecuador | Orellana | Yasuní Scientific Station | This work |
Ecuador | Orellana | Yuturi | MECN 1517 |
Ecuador | Orellana | Pindo, bloque 64 | MZUTI 3957 |
Ecuador | Pastaza | Arutam, 5 km S of | iNaturalist |
Ecuador | Pastaza | Balsaura | Avila-Pires 1995 |
Ecuador | Pastaza | Balsaura | Ortega-Andrade 2010 |
Ecuador | Pastaza | Between Sarayacu and Canelos | Ribeiro-Júnior 2015 |
Ecuador | Pastaza | Conambo | Ortega-Andrade 2010 |
Ecuador | Pastaza | Destacamento Militar Shiona | Avila-Pires 1995 |
Ecuador | Pastaza | Juyuintza | Ortega-Andrade 2010 |
Ecuador | Pastaza | Kallana | MZUTI 5087 |
Ecuador | Pastaza | Kapawi Lodge | iNaturalist |
Ecuador | Pastaza | Lorocachi | Torres-Carvajal et al. 2019 |
Ecuador | Pastaza | Palanda | Ribeiro-Júnior 2015 |
Ecuador | Pastaza | Puerto Murialdo | MECN 3821 |
Ecuador | Pastaza | Puyo, nearby | KU 178947 |
Ecuador | Pastaza | Quebrada Churoyacu | Ribeiro-Júnior 2015 |
Ecuador | Pastaza | Río Bufeo | Ortega-Andrade 2010 |
Ecuador | Pastaza | Río Lliquino | Ribeiro-Júnior 2015 |
Ecuador | Pastaza | Sarayacu | This work |
Ecuador | Pastaza | UNOCAL Base Camp | Ribeiro-Júnior 2015 |
Ecuador | Pastaza | Zanja Arajuno | This work |
Ecuador | Sucumbíos | Blanca | iNaturalist |
Ecuador | Sucumbíos | Cuyabeno Lodge | iNaturalist |
Ecuador | Sucumbíos | Cuyabeno River Lodge | iNaturalist |
Ecuador | Sucumbíos | Dureno | Duellman 1978 |
Ecuador | Sucumbíos | Estación Amazonas OCP | Valencia & Garzón 2011 |
Ecuador | Sucumbíos | Garzacocha | iNaturalist |
Ecuador | Sucumbíos | La Selva Lodge | iNaturalist |
Ecuador | Sucumbíos | Lago Agrio | Duellman 1978 |
Ecuador | Sucumbíos | Laguna de Pañacocha | iNaturalist |
Ecuador | Sucumbíos | Limoncocha | D’Angiolella et al. 2011 |
Ecuador | Sucumbíos | Lower Río Cuyabeno | iNaturalist |
Ecuador | Sucumbíos | Mocoa | iNaturalist |
Ecuador | Sucumbíos | Parque Perla | MZUTI 2561 |
Ecuador | Sucumbíos | Pozo Tetete | iNaturalist |
Ecuador | Sucumbíos | Precooperativa Centinela | Alemán 2013 |
Ecuador | Sucumbíos | Puerto Ore | Duellman 1978 |
Ecuador | Sucumbíos | Remolino | iNaturalist |
Ecuador | Sucumbíos | Reserva Chichicos | Photo by Guido Bladimir |
Ecuador | Sucumbíos | Reserva Ecológica Cofán Bermejo | MECN 8290 |
Ecuador | Sucumbíos | Río Blanco | iNaturalist |
Ecuador | Sucumbíos | San Pablo de Kantesiya | MECN 390 |
Ecuador | Sucumbíos | Sani Lodge | iNaturalist |
Ecuador | Sucumbíos | Sani Lodge | This work |
Ecuador | Sucumbíos | Santa Cecilia | D’Angiolella et al. 2011 |
Ecuador | Sucumbíos | Shushufindi, 4 km N of | iNaturalist |
Ecuador | Sucumbíos | Waita Amazon Lodge | iNaturalist |
Ecuador | Sucumbíos | Zancudococha | Felipe Campos, pers. comm. |
Perú | Amazonas | Aintami | Ribeiro-Júnior 2015 |
Perú | Amazonas | Boca del Río Santiago | D’Angiolella et al. 2011 |
Perú | Amazonas | Chigkan Entse | Ribeiro-Júnior 2015 |
Perú | Amazonas | Huampami | Ribeiro-Júnior 2015 |
Perú | Amazonas | Kayamas | Ribeiro-Júnior 2015 |
Perú | Amazonas | La Poza | Ribeiro-Júnior 2015 |
Perú | Amazonas | Mouth of Kagka river | USNM 316699 |
Perú | Amazonas | Puerto Galilea | Ribeiro-Júnior 2015 |
Perú | Amazonas | Quebrada Sasa | Ribeiro-Júnior 2015 |
Perú | Amazonas | Quebrada Shinganatza | Almendáriz et al. 2014 |
Perú | Amazonas | Río Caterpiza | Ribeiro-Júnior 2015 |
Perú | Amazonas | Río Cenepa | Ribeiro-Júnior 2015 |
Perú | Amazonas | San Antonio, Río Cenepa | Ribeiro-Júnior 2015 |
Perú | Amazonas | Shiringa | Ribeiro-Júnior 2015 |
Perú | Amazonas | Yutupis | Ribeiro-Júnior 2015 |
Perú | Loreto | Barranca | Ribeiro-Júnior 2015 |
Perú | Loreto | Estación Biológica Pithecia | Avila-Pires 1995 |
Perú | Loreto | Galicia | D’Angiolella et al. 2011 |
Perú | Loreto | Güeppí | FMNH 2008 |
Perú | Loreto | Pongo Chinim | FMNH 2012 |
Perú | Loreto | Río Pacaya | D’Angiolella et al. 2011 |
Perú | Loreto | Río Yuvineto | MNHN 1978.2364 |