Published May 16, 2023. Open access.

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Slender Anole (Anolis fuscoauratus)

Reptiles of Ecuador | Sauria | Anolidae | Anolis fuscoauratus

English common names: Slender Anole, Slender Amazon Anole, Brown-eared Anole.

Spanish common names: Anolis esbelto, anolis delgado.

Recognition: ♂♂ 15.2 cmMaximum distance from the snout to the tip of the tail. Snout–vent length=5.1 cm. ♀♀ 15.6 cmMaximum distance from the snout to the tip of the tail. Snout–vent length=5.4 cm..1,2 Anoles are easily distinguishable from other lizards by their diurnal habits, extensible dewlap in males, expanded digital pads, and granular scales on the dorsum and belly. The Slender Anole (Anolis fuscoauratus) can be identified based on its small body size, uniform brown coloration, and rose-pink dewlap with white scales in males.3,4 Females usually have a pale cream vertebral stripe (Fig. 1) and a rudimentary dewlap.3,5 Anolis fuscoauratus is often confused with A. ortonii, which has a yellow-striped bright orange dewlap, and A. trachyderma, which has a small orange-yellow dewlap with black scales.6,7

Figure showing variation among individuals of Anolis fuscoauratus

Figure 1: Individuals of Anolis fuscoauratus from Maycu Reserve, Zamora Chinchipe province, Ecuador.

Natural history: Anolis fuscoauratus is usually the most common anole wherever it occurs,2,8,9 particularly in terra firme forest, although it also occurs in igapó (=blackwater-flooded forests) and inundated forests along white-water rivers.7,10,11 This species inhabits pristine rainforests as well as logged forests, edge situations, tree plantations, banana groves, cacao groves, and rural gardens.26 Slender Anoles are included in the “grass-bush” anole ecomorph12 because they primarily use the undergrowth and herbaceous forest strata.7,9 However, individuals foraging on branches at 35 m above the ground are not uncommon.2 Activity occurs from sunrise to sunset,11 especially on tree trunks and palm fronds,13 but also on twigs, branches, banana leaves, logs, leaf-litter, shrubs, small saplings, and fence posts within 1 m from the ground.314 Slender Anoles are primarily active in the shade,11 at ambient temperatures ~27°C,11 but also bask during the early morning.47 At night, individuals roost at heights of 0.1–8.4 m above the ground on flimsy perches (mostly leaves, twigs, and grasses)1,6,7 that will move if disturbed by a predator.2 Anolis fuscoauratus is an ambush predator7,9 that feeds primarily on spiders, beetles, and crickets,11,13 but also on at least 20 other prey items, from insect larvae to stingless bees and pseudoscorpions.68 Males defend territories by performing head bobs and dewlap displays.7 Females lay one egg at a time,9 up to three times per year,15 and the incubation period is 53–63 days (~2 months).6 Breeding seems to take place year-round throughout the Amazon.4 Slender Anoles escape predators by remaining perfectly still; which, combined with their twig camouflage, makes them hard to detect.2 If on a trunk, they move to the opposite side or run up along the trunk.4,8,9 Predators include snakes (notably Imantodes cenchoa,7,16 but also Chironius multiventris,17 Drymoluber dichrous,18 and Xenoxybelis argenteus4), birds (Monasa morphoeus),2 whiptail lizards,19 spiders,20 and whipscorpions.21 Roaches have been observed feeding on the tears of an A. fuscoauratus.22

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Conservation: Least Concern Believed to be safe from extinction given current circumstances..23 Anolis fuscoauratus is listed in this category given its wide distribution, presence in major protected areas, lack of widespread threats, and presumed large stable populations.23

Distribution: Anolis fuscoauratus is widely distributed throughout the Amazon basin in Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname, and Venezuela.10 The species also occurs in the Atlantic Forest of Brazil.10 In Ecuador, A. fuscoauratus has been recorded at elevations between 99 and 1920 m (Fig. 2).

Distribution of Anolis fuscoauratus in Ecuador

Figure 2: Distribution of Anolis fuscoauratus in Ecuador. See Appendix 1 for a complete list of the presence localities included in the map.

Etymology: The generic name Anolis is thought to have originated from Cariban languages, specifically from the word anoli, which is the name Arawak peoples may have used to refer to this group of lizards.24 The specific epithet fuscoauratus comes from the Latin words fuscus (=dusky) and auratus (=golden),25 and refers to the coloration of the alcohol-preserved holotype.

See it in the wild: Slender Anoles are easily located in forested areas throughout the species’ area of distribution in Ecuador. Some of the best localities to find these lizards are Yasuní Scientific Station, Cuyabeno Reserve, and Jatun Sacha Biological Reserve. The anoles can be spotted by scanning the understory vegetation along forest trails at night.

Author: Alejandro ArteagaaAffiliation: Khamai Foundation, Quito, Ecuador.

Photographer: Jose VieirabAffiliation: Tropical Herping (TH), Quito, Ecuador.,cAffiliation: ExSitu, Quito, Ecuador.

How to cite? Arteaga A (2023) Slender Anole (Anolis fuscoauratus). In: Arteaga A, Bustamante L, Vieira J (Eds) Reptiles of Ecuador: Life in the middle of the world. Available from: DOI: 10.47051/XDPY7871

Literature cited:

  1. Ayala-Varela F (2004) Revisión taxonómica y de variación geográfica de las especies de Anolis (Sauria: Polychrotidae) del Oriente Ecuatoriano. BSc thesis, Pontificia Universidad Católica del Ecuador, 204 pp.
  2. Field notes, Reptiles of Ecuador book project.
  3. Castro Herrera F, Ayala SC (1988) Saurios de Colombia. Unpublished, Bogotá, 692 pp.
  4. Avila-Pires TCS (1995) Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen 299: 1–706.
  5. Dixon JR, Soini P (1986) The reptiles of the upper Amazon Basin, Iquitos region, Peru. Milwaukee Public Museum, Milwaukee, 154 pp.
  6. Duellman WE (1978) The biology of an equatorial herpetofauna in Amazonian Ecuador. Publications of the Museum of Natural History, University of Kansas 65: 1–352.
  7. Duellman WE (2005) Cusco amazónico: the lives of amphibians and reptiles in an Amazonian rainforest. Cornell University Press, Ithaca, 433 pp.
  8. Beebe W (1944) Field notes on the lizards of Kartabo, British Guiana, and Caripito, Venezuela. Part 2. Iguanidae. Zoologica 29: 195–216.
  9. Vitt LJ, De la Torre S (1996) A research guide to the lizards of Cuyabeno. Pontificia Universidad Católica del Ecuador, Quito, 165 pp.
  10. Ribeiro-Junior MA (2015) Catalogue of distribution of lizards (Reptilia: Squamata) from the Brazilian Amazonia. I. Dactyloidae, Hoplocercidae, Iguanidae, Leiosauridae, Polychrotidae, Tropiduridae. Zootaxa 3983: 001–110. DOI: 10.11646/zootaxa.3983.1.1
  11. Vitt LJ, Avila-Pires TCS, Zani PA, Sartorius SS, Espósito MC (2003) Life above ground: ecology of Anolis fuscoauratus in the Amazon rain forest, and comparisons with its nearest relatives. Canadian Journal of Zoology 81: 142–156. DOI: 10.1139/z02-230
  12. Moreno-Arias R, Velasco JA, Urbina Cardona J, Cárdenas-Arévalo G, Medina Rangel G, Gutiérrez Cárdenas P, Olaya-Rodriguez M, Noguera-Urbano E (2021) Atlas de la biodiversidad de Colombia. Anolis. Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, Bogotá, 72 pp.
  13. Pinto Aguirre JA (2014) Ecología de una comunidad de lagartijas del género Anolis en el Parque Nacional Yasuní. BSc thesis, Quito, Pontificia Universidad Católica del Ecuador, 52 pp.
  14. Vitt LJ, Zani PA (1996) Organization of a taxonomically diverse lizard assemblage in Amazonian Ecuador. Canadian Journal of Zoology 74: 1313–1335.
  15. Vitt LJ, Magnusson WE, Avila-Pires TCS, Pimentel Lima A (2008) Guide to the lizards of Reserva Adolpho Ducke, Central Amazonia. Áttema Design Editorial, Manaus, 176 pp.
  16. Martins M, Oliveira ME (1998) Natural history of snakes in forests of the Manaus region, Central Amazonia, Brazil. Herpetological Natural History 6: 78–150.
  17. Roberto IJ, Ramos Souza A (2020) Review of prey items recorded for snakes of the genus Chironius (Squamata, Colubridae), including the first record of Osteocephalus as prey. Herpetology Notes 13: 1–5.
  18. Veríssimo D, de Freitas MA, Rosa GM (2012) Drymoluber dichrous (Northern Woodland Racer) and Anolis fuscoauratus (Slender Anole): predation. Herpetological Bulletin 120: 36–38.
  19. Franzini LD, Martins Teixeira AA, Alves Teles D, de Araujo Filho JA, Oliveira Mesquita D (2017) Predation of Norops fuscoauratus (Duméril and Bibron, 1837) by Kentropyx calcarata (Spix, 1825) in a remnant of Atlantic Forest, Brazil. Herpetology Notes 10: 249–250.
  20. Reyes-Olivares C, Guajardo-Santibáñez A, Segura B, Zañartu N, Penna M, Labra A (2020) Lizard predation by spiders: a review from the Neotropical and Andean regions. Ecology and Evolution 10: 10953–10964. DOI: 10.1002/ece3.6801
  21. Mendes DMM, Barros AL, Pires DA, Viana PF (2017) Anolis fuscoauratus (Slender Anole): predation. Herpetological Review 48: 635–636.
  22. van den Burg MP, González de Rueda JA (2021) Lachryphagy by cockroaches: reptile tears to increase reproductive output? Neotropical Biodiversity 7: 276–278. DOI: 10.1080/23766808.2021.1953892
  23. Ibáñez R, Jaramillo C, Köhler G, Gutiérrez-Cárdenas P, Rivas G, Cisneros-Heredia DF, Perez P, Aparicio J, Avila-Pires TCS, Gonzales L (2014) Anolis fuscoauratus. The IUCN Red List of threatened species. Available from: DOI: 10.2305/IUCN.UK.2020-3.RLTS.T203091A2760203.en
  24. Allsopp R (1996) Dictionary of Caribbean English Usage. Oxford University Press, Oxford, 776 pp.
  25. Brown RW (1956) Composition of scientific words. Smithsonian Books, Washington D.C., 882 pp.

Appendix 1: Locality data used to create the distribution map of Anolis fuscoauratus in Ecuador (Fig. 2). Go to the section on symbols and abbreviations for a list of acronyms used.

ColombiaAmazonasPuerto NariñoRibeiro-Júnior & Amaral 2016
ColombiaCaquetáCarmelo, 4 km S ofGutiérrez-Lamus et al. 2020
ColombiaCaquetáCarmelo, 7.4 km S ofGutiérrez-Lamus et al. 2020
ColombiaCaquetáIsla ViejaRibeiro-Júnior & Amaral 2016
ColombiaCaquetáLarandiaiNaturalist; photo examined
ColombiaCaquetáLos AlicangarosMCZ 118133; Harvard University & Morris 2023
ColombiaPutumayoCentro Experimental AmazónicoiNaturalist; photo examined
ColombiaPutumayoEscuela Santa TeresaiNaturalist; photo examined
ColombiaPutumayoFinca CharapaIAvH-R-4870; Borja-Acosta & Galeano Muñoz 2023
ColombiaPutumayoLas CameliasCahueño & Barbosa 2022
ColombiaPutumayoMocoa, 10 km S ofRibeiro-Júnior & Amaral 2016
ColombiaPutumayoNariñoGeopark Colombia 2022
ColombiaPutumayoReserva La Isla EscondidaiNaturalist; photo examined
ColombiaPutumayoSan PedroiNaturalist; photo examined
ColombiaPutumayoSanta Ana campRibeiro-Júnior & Amaral 2016
ColombiaPutumayoSimón BolívarIAvH-R-9157; Borja-Acosta & Ocampo 2021
ColombiaPutumayoVereda Islas de CartagenaIAvH-R-9151; Borja-Acosta & Galeano Muñoz 2023
ColombiaPutumayoVereda Monte VerdeIAvH-R-9158; Borja-Acosta & Galeano Muñoz 2023
ColombiaPutumayoVereda PeneyaIAvH-CT-39150; Borja-Acosta & Galeano Muñoz 2023
ColombiaPutumayoVillagarzón, 12 km SW ofICN 4452; Calderón et al. 2023
EcuadorAzuaySopladoraMZUA.RE.0222; examined
EcuadorMorona SantiagoCamino Gualaquiza–ZamoraAyala-Varela 2004
EcuadorMorona SantiagoChiguazaRibeiro-Júnior & Amaral 2016
EcuadorMorona SantiagoCusuimeOrtega-Andrade 2010
EcuadorMorona SantiagoEl TiinkReptiles of Ecuador book database
EcuadorMorona SantiagoEstación Biológica WisuiChaparro et al. 2011
EcuadorMorona SantiagoJimbitonoAyala-Varela 2004
EcuadorMorona SantiagoMacasRibeiro-Júnior & Amaral 2016
EcuadorMorona SantiagoMacumaiNaturalist; photo examined
EcuadorMorona SantiagoNuevo IsraelQCAZ 10085; Ayala-Varela & Carvajal-Campos 2022
EcuadorMorona SantiagoParque Nacional SangayBrito & Almendariz 2013
EcuadorMorona SantiagoSan JorgeMZUTI 2872; examined
EcuadorMorona SantiagoSan Juan BoscoMZUA.RE.0179; examined
EcuadorMorona SantiagoSawastianOrtega-Andrade 2010
EcuadorMorona SantiagoShuin MamusiNaturalist; photo examined
EcuadorMorona SantiagoSiete Iglesias ReserveOnline multimedia
EcuadorMorona SantiagoSucúaRibeiro-Júnior & Amaral 2016
EcuadorMorona SantiagoTaishaRibeiro-Júnior & Amaral 2016
EcuadorMorona SantiagoTres RanchosiNaturalist; photo examined
EcuadorMorona SantiagoTukupiQCAZ 4939; Ayala-Varela & Carvajal-Campos 2022
EcuadorMorona SantiagoTurulaOrtega-Andrade 2010
EcuadorMorona SantiagoVía Peñas-ShaimiQCAZ 16652; Ayala-Varela & Carvajal-Campos 2022
EcuadorMorona SantiagoVilla AshuaraOrtega-Andrade 2010
EcuadorNapoAnaconda LodgeOnline multimedia
EcuadorNapoArchidonaRibeiro-Júnior & Amaral 2016
EcuadorNapoBaezaRibeiro-Júnior & Amaral 2016
EcuadorNapoEl ChacoMHNG 2464.026; not examined
EcuadorNapoGareno LodgeReptiles of Ecuador book database
EcuadorNapoHamadryade LodgeReptiles of Ecuador book database
EcuadorNapoHidroeléctrica Coca Codo SinclairCOCASINCLAIR 2013
EcuadorNapoHollín–Loreto roadReptiles of Ecuador book database
EcuadorNapoIkiamPhoto by Grace Reyes
EcuadorNapoJatun Sacha Biological StationRibeiro-Júnior & Amaral 2016
EcuadorNapoLiana LodgeReptiles of Ecuador book database
EcuadorNapoMisahuallíRibeiro-Júnior & Amaral 2016
EcuadorNapoNarupa ReserveReptiles of Ecuador book database
EcuadorNapoPacto SumacoMZUTI 5317; examined
EcuadorNapoRío PayaminoRibeiro-Júnior & Amaral 2016
EcuadorNapoSan Jose Viejo de SumacoRibeiro-Júnior & Amaral 2016
EcuadorNapoSan RafaelRibeiro-Júnior & Amaral 2016
EcuadorNapoSinchi SachaPhoto by Ernesto Arbeláez
EcuadorNapoSuchipakariReptiles of Ecuador book database
EcuadorNapoWild Sumaco Wildlife SanctuaryCamper et al. 2021
EcuadorNapoYachana LodgeReptiles of Ecuador book database
EcuadorNapoYachana ReserveBeirne et al. 2013
EcuadorOrellanaBigayKingsbury et al. 2008
EcuadorOrellanaBloque 31Libro PetroAmazonas
EcuadorOrellanaBloque 65MZUTI 3728; not examined
EcuadorOrellanaComunidad El EdénAyala-Varela 2004
EcuadorOrellanaCordillera GalerasAyala-Varela 2004
EcuadorOrellanaEl CocaRibeiro-Júnior & Amaral 2016
EcuadorOrellanaHacienda PrimaveraRibeiro-Júnior & Amaral 2016
EcuadorOrellanaLa Joya de los SachasMHNG 2463.085; not examined
EcuadorOrellanaLoretoRibeiro-Júnior & Amaral 2016
EcuadorOrellanaMandaripanga LodgeReptiles of Ecuador book database
EcuadorOrellanaMouth of Río GuataracoRibeiro-Júnior & Amaral 2016
EcuadorOrellanaNapo Wildlife CenterReptiles of Ecuador book database
EcuadorOrellanaNenkepareReptiles of Ecuador book database
EcuadorOrellanaPacca HuasiKingsbury et al. 2008
EcuadorOrellanaPaushiyacuKingsbury et al. 2008
EcuadorOrellanaPindoiNaturalist; photo examined
EcuadorOrellanaRío AguaricoTapia-del Águila et al. 2019
EcuadorOrellanaRío BigalGarcía et al. 2021
EcuadorOrellanaRío NashiñoQCAZ 5493; Ayala-Varela & Carvajal-Campos 2022
EcuadorOrellanaRío YasuníQCAZ 4778; Ayala-Varela & Carvajal-Campos 2022
EcuadorOrellanaRío Yasuní, near Lake JatuncochaReptiles of Ecuador book database
EcuadorOrellanaRosa FloridaQCAZ 6055; Ayala-Varela & Carvajal-Campos 2022
EcuadorOrellanaSan José de PayaminoMaynard et al. 2016
EcuadorOrellanaShiripuno LodgeiNaturalist; photo examined
EcuadorOrellanaTiputini Biodiversity Station Cisneros-Heredia 2003
EcuadorOrellanaYasuni Scientific StationReptiles of Ecuador book database
EcuadorPastazaAlto CurarayRibeiro-Júnior & Amaral 2016
EcuadorPastazaArajuno Ribeiro-Júnior & Amaral 2016
EcuadorPastazaBalsauraOrtega-Andrade 2010
EcuadorPastazaBataburo LodgeOrtega-Andrade 2010
EcuadorPastazaCabañas PiatúaReptiles of Ecuador book database
EcuadorPastazaCabeceras del Río BobonazaReptiles of Ecuador book database
EcuadorPastazaChichirotaRibeiro-Júnior & Amaral 2016
EcuadorPastazaChontoaRibeiro-Júnior & Amaral 2016
EcuadorPastazaConamboOrtega-Andrade 2010
EcuadorPastazaCopatazaRibeiro-Júnior & Amaral 2016
EcuadorPastazaCuraray MedioReptiles of Ecuador book database
EcuadorPastazaDon TomásRibeiro-Júnior & Amaral 2016
EcuadorPastazaEntre Sarayacu y CanelosRibeiro-Júnior & Amaral 2016
EcuadorPastazaEstación Agro-Ecológica FátimaProaño-Morales et al. 2017
EcuadorPastazaFinca HeimatlosPhoto by Ferhat Gundogdu
EcuadorPastazaGuacheOrtega-Andrade 2010
EcuadorPastazaIndichurisiNaturalist; photo examined
EcuadorPastazaJuyuintzaOrtega-Andrade 2010
EcuadorPastazaKapawi LodgeReptiles of Ecuador book database
EcuadorPastazaKurintzaOrtega-Andrade 2010
EcuadorPastazaLlanchamacochaAyala-Varela 2004
EcuadorPastazaLorocachiQCAZ 11722; Ayala-Varela & Carvajal-Campos 2022
EcuadorPastazaMeraRibeiro-Júnior & Amaral 2016
EcuadorPastazaMontalvoOrtega-Andrade 2010
EcuadorPastazaMoreteRibeiro-Júnior & Amaral 2016
EcuadorPastazaMouth of Río ShionayacuRibeiro-Júnior & Amaral 2016
EcuadorPastazaNuevo GolondrinaRibeiro-Júnior & Amaral 2016
EcuadorPastazaPalandaRibeiro-Júnior & Amaral 2016
EcuadorPastazaPeter Archer’s placePhoto by Yatin Kalki
EcuadorPastazaPindoyacuOrtega-Andrade 2010
EcuadorPastazaPozo Petrolero MisiónAlmendáriz 1987
EcuadorPastazaPuyoRibeiro-Júnior & Amaral 2016
EcuadorPastazaRío AlpayacuRibeiro-Júnior & Amaral 2016
EcuadorPastazaRío Anzu ReserveReyes-Puig & Tobar 2008
EcuadorPastazaRío Arajuno, headwaters ofRibeiro-Júnior & Amaral 2016
EcuadorPastazaRío BufeoOrtega-Andrade 2010
EcuadorPastazaRío CorrientesRibeiro-Júnior & Amaral 2016
EcuadorPastazaRío LliquinoRibeiro-Júnior & Amaral 2016
EcuadorPastazaRío PindoRibeiro-Júnior & Amaral 2016
EcuadorPastazaRío RutunoRibeiro-Júnior & Amaral 2016
EcuadorPastazaRío VillanoRibeiro-Júnior & Amaral 2016
EcuadorPastazaSarayacuRibeiro-Júnior & Amaral 2016
EcuadorPastazaShellRibeiro-Júnior & Amaral 2016
EcuadorPastazaSumak Kawsay In SituBentley et al. 2021
EcuadorPastazaTamandúa LodgePhoto by Jorge Flores
EcuadorPastazaTigüinoOrtega-Andrade 2010
EcuadorPastazaUNOCAL Base CampRibeiro-Júnior & Amaral 2016
EcuadorPastazaVeracruzRibeiro-Júnior & Amaral 2016
EcuadorPastazaZanja ArajunoReptiles of Ecuador book database
EcuadorSucumbíosBloque 27, Sector Blanca ADHMECN 8334; not examined
EcuadorSucumbíosCabañas Neotropic TurisAyala-Varela 2004
EcuadorSucumbíosCaimán LodgeReptiles of Ecuador book database
EcuadorSucumbíosCampo PlatanilloEnvirotec 2015
EcuadorSucumbíosComunidad Secoya RemolinoiNaturalist; photo examined
EcuadorSucumbíosComunidad ZábaloCevallos Bustos 2010
EcuadorSucumbíosDurenoRibeiro-Júnior & Amaral 2016
EcuadorSucumbíosEl ReventadorMHNG 2275.087; not examined
EcuadorSucumbíosEstación PUCE CuyabenoAyala-Varela 2004
EcuadorSucumbíosGarzacochaYánez-Muñoz & Venegas 2008
EcuadorSucumbíosGüeppicilloYánez-Muñoz & Venegas 2008
EcuadorSucumbíosInecel StationRibeiro-Júnior & Amaral 2016
EcuadorSucumbíosLa BalsareñaiNaturalist; photo examined
EcuadorSucumbíosLa Selva LodgeReptiles of Ecuador book database
EcuadorSucumbíosLago AgrioDuellman 1978
EcuadorSucumbíosLaguna PañacochaMHNG 2590.073; not examined
EcuadorSucumbíosLimoncocha Biological ReserveRibeiro-Júnior & Amaral 2016
EcuadorSucumbíosLumbaquiReyes-Puig & Tobar 2008
EcuadorSucumbíosNear Río CuyabenoVitt et al. 2003
EcuadorSucumbíosPetroamazonas campReptiles of Ecuador book database
EcuadorSucumbíosPlataforma Espejo 1Consultora Cinge 2012
EcuadorSucumbíosPlayas del CuyabenoUIMNH 64332; not examined
EcuadorSucumbíosPuerto LibreDuellman 1978
EcuadorSucumbíosPuerto OreDuellman 1978
EcuadorSucumbíosRío Piedra FinaiNaturalist; photo examined
EcuadorSucumbíosRío SingueiNaturalist; photo examined
EcuadorSucumbíosSacha LodgePhoto by Ameet Zaveri
EcuadorSucumbíosSan Pablo de KantesiyaRibeiro-Júnior & Amaral 2016
EcuadorSucumbíosSani LodgeReptiles of Ecuador book database
EcuadorSucumbíosSanta Cecilia Duellman 1978
EcuadorSucumbíosShushufindiRibeiro-Júnior & Amaral 2016
EcuadorSucumbíosTarapoaMHNG 2460.01; not examined
EcuadorSucumbíosWaita Amazon LodgeiNaturalist; photo examined
EcuadorTungurahuaAlong Ulba riverReptiles of Ecuador book database
EcuadorTungurahuaReserva La CandelariaReptiles of Ecuador book database
EcuadorTungurahuaRío NegroAyala-Varela 2004
EcuadorTungurahuaRío VerdeAyala-Varela 2004
EcuadorZamora ChinchipeCopalingaReeves et al.
EcuadorZamora ChinchipeEstación Experimental El PadmiGonzález et al. 2010
EcuadorZamora ChinchipeFina PermatreeiNaturalist; photo examined
EcuadorZamora ChinchipeReserva MaycuReptiles of Ecuador book database
EcuadorZamora ChinchipeReserva Palmitera ValladolidReptiles of Ecuador book database
EcuadorZamora ChinchipeTepuy Las OrquídeasReptiles of Ecuador book database
EcuadorZamora ChinchipeTunantzaiNaturalist; photo examined
EcuadorZamora ChinchipeValle del QuimiBetancourt et al. 2018
EcuadorZamora ChinchipeYacuri–ZumbaReptiles of Ecuador book database
EcuadorZamora ChinchipeYankuam LodgeiNaturalist; photo examined
EcuadorZamora ChinchipeZamoraAyala-Varela 2004
EcuadorZamora ChinchipeZumba–Pucubamba roadReptiles of Ecuador book database
EcuadorZamora ChinchipeZurmiReptiles of Ecuador book database
PeruAmazonasAlfonso UgarteRibeiro-Júnior & Amaral 2016
PeruAmazonasBoca del Río SantiagoRibeiro-Júnior & Amaral 2016
PeruAmazonasCaterpizaRibeiro-Júnior & Amaral 2016
PeruAmazonasFalso PaquishaAlmendáriz et al. 2014
PeruAmazonasHuampamiRibeiro-Júnior & Amaral 2016
PeruAmazonasLa PozaRibeiro-Júnior & Amaral 2016
PeruAmazonasPaagatRibeiro-Júnior & Amaral 2016
PeruAmazonasPastazilloRibeiro-Júnior & Amaral 2016
PeruAmazonasPuerto GalileaRibeiro-Júnior & Amaral 2016
PeruAmazonasQuebrada SasaRibeiro-Júnior & Amaral 2016
PeruAmazonasQuebrada ShinganatzaAlmendáriz et al. 2014
PeruAmazonasSan Antonio, Río CenepaRibeiro-Júnior & Amaral 2016
PeruAmazonasShaimRibeiro-Júnior & Amaral 2016
PeruAmazonasShiringaRibeiro-Júnior & Amaral 2016
PeruAmazonasTseasimRibeiro-Júnior & Amaral 2016
PeruAmazonasYutupisRibeiro-Júnior & Amaral 2016
PeruLoretoAguas NegrasYánez-Muñoz & Venegas 2008
PeruLoretoCampo AndoasValqui Schult 2015
PeruLoretoCentro UniónRibeiro-Júnior & Amaral 2016
PeruLoretoCerro de KampankisCatenazzi & Venegas 2016
PeruLoretoCocama LodgeiNaturalist; photo examined
PeruLoretoEstirónRibeiro-Júnior & Amaral 2016
PeruLoretoLagartocochaKnell 2012
PeruLoretoMishanaRibeiro-Júnior & Amaral 2016
PeruLoretoRedondocochaYánez-Muñoz & Venegas 2008
PeruLoretoRío Lagarto Cocha, Aguas NegrasRibeiro-Júnior & Amaral 2016
PeruLoretoSuno LlactaRibeiro-Júnior & Amaral 2016
PeruLoretoZona Reservada GüeppíYánez-Muñoz & Venegas 2008